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The Origin of Species: Endnotes

The Origin of Species
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table of contents
  1. Titlepage
  2. Imprint
  3. Epigraphs
  4. An Historical Sketch of the Progress of Opinion on the Origin of Species, Previously to the Publication of the First Edition of This Work
  5. Introduction
  6. The Origin of Species by Means of Natural Selection
    1. I: Variation Under Domestication
      1. Causes of Variability
      2. Effects of Habit and of the Use or Disuse of Parts; Correlated Variation; Inheritance
      3. Character of Domestic Varieties; Difficulty of Distinguishing Between Varieties and Species; Origin of Domestic Varieties from One or More Species
      4. Breeds of the Domestic Pigeon, Their Differences and Origin
      5. Principles of Selection Anciently Followed, and Their Effects
      6. Unconscious Selection
      7. Circumstances Favourable to Man’s Power of Selection
    2. II: Variation Under Nature
      1. Individual Differences
      2. Doubtful Species
      3. Wide-Ranging, Much Diffused, and Common Species Vary Most
      4. Species of the Larger Genera in Each Country Vary More Frequently Than the Species of the Smaller Genera
      5. Many of the Species Included Within the Larger Genera Resemble Varieties in Being Very Closely, but Unequally, Related to Each Other, and in Having Restricted Ranges
      6. Summary
    3. III: Struggle for Existence
      1. The Term, Struggle for Existence, Used in a Large Sense
      2. Geometrical Ratio of Increase
      3. Nature of the Checks to Increase
      4. Complex Relations of All Animals and Plants to Each Other in the Struggle for Existence
      5. Struggle for Life Most Severe Between Individuals and Varieties of the Same Species
    4. IV: Natural Selection; or the Survival of the Fittest
      1. Sexual Selection
      2. Illustrations of the Action of Natural Selection, or the Survival of the Fittest
      3. On the Intercrossing of Individuals
      4. Circumstances Favourable for the Production of New Forms Through Natural Selection
      5. Extinction Caused by Natural Selection
      6. Divergence of Character
      7. The Probable Effects of the Action of Natural Selection Through Divergence of Character and Extinction, on the Descendants of a Common Ancestor
      8. On the Degree to Which Organisation Tends to Advance
      9. Convergence of Character
      10. Summary of Chapter
    5. V: Laws of Variation
      1. Effects of the Increased Use and Disuse of Parts, as Controlled by Natural Selection
      2. Acclimatisation
      3. Correlated Variation
      4. Compensation and Economy of Growth
      5. Multiple, Rudimentary, and Lowly-Organised Structures Are Variable
      6. A Part Developed in Any Species in an Extraordinary Degree or Manner, in Comparison with the Same Part in Allied Species, Tends to Be Highly Variable
      7. Specific Characters More Variable Than Generic Characters
      8. Secondary Sexual Characters Variable
      9. Distinct Species Present Analogous Variations, So That a Variety of One Species Often Assumes a Character Proper to an Allied Species, or Reverts to Some of the Characters of an Early Progenitor
      10. Summary
    6. VI: Difficulties of the Theory
      1. On the Absence or Rarity of Transitional Varieties
      2. On the Origin and Transition of Organic Beings with Peculiar Habits and Structure
      3. Organs of Extreme Perfection and Complication
      4. Modes of Transition
      5. Special Difficulties of the Theory of Natural Selection
      6. Organs of Little Apparent Importance, as Affected by Natural Selection
      7. Utilitarian Doctrine, How Far True: Beauty, How Acquired
      8. Summary: The Law of Unity of Type and of the Conditions of Existence Embraced by the Theory of Natural Selection
    7. VII: Miscellaneous Objections to the Theory of Natural Selection
    8. VIII: Instinct
      1. Inherited Changes of Habit or Instinct in Domesticated Animals
      2. Special Instincts
      3. Instincts of the Cuckoo
      4. Slave-Making Instinct
      5. Cell-Making Instinct of the Hive-Bee
      6. Objections to the Theory of Natural Selection as Applied to Instincts: Neuter and Sterile Insects
      7. Summary
    9. IX: Hybridism
      1. Degrees of Sterility
      2. Laws Governing the Sterility of First Crosses and of Hybrids
      3. Origin and Causes of the Sterility of First Crosses and of Hybrids
      4. Reciprocal Dimorphism and Trimorphism
      5. Fertility of Varieties When Crossed, and of Their Mongrel Offspring, Not Universal
      6. Hybrids and Mongrels Compared, Independently of Their Fertility
      7. Summary of Chapter
    10. X: On the Imperfection of the Geological Record
      1. On the Lapse of Time, as Inferred from the Rate of Deposition and Extent of Denudation
      2. On the Poorness of Palaeontological Collections
      3. On the Absence of Numerous Intermediate Varieties in Any Single Formation
      4. On the Sudden Appearance of Whole Groups of Allied Species
      5. On the Sudden Appearance of Groups of Allied Species in the Lowest Known Fossiliferous Strata
    11. XI: On the Geological Succession of Organic Beings
      1. On Extinction
      2. On the Forms of Life Changing Almost Simultaneously Throughout the World
      3. On the Affinities of Extinct Species to Each Other, and to Living Forms
      4. On the State of Development of Ancient Compared with Living Forms
      5. On the Succession of the Same Types Within the Same Areas, During the Later Tertiary Periods
      6. Summary of the Preceding and Present Chapters
    12. XII: Geographical Distribution
      1. Single Centres of Supposed Creation
      2. Means of Dispersal
      3. Dispersal During the Glacial Period
      4. Alternate Glacial Periods in the North and South
    13. XIII: Geographical Distribution—Continued
      1. Freshwater Productions
      2. On the Inhabitants of Oceanic Islands
      3. Absence of Batrachians and Terrestrial Mammals on Oceanic Islands
      4. On the Relations of the Inhabitants of Islands to Those of the Nearest Mainland
      5. Summary of the Last and Present Chapters
    14. XIV: Mutual Affinities of Organic Beings: Morphology—Embryology—Rudimentary Organs—Classification
      1. Analogical Resemblances
      2. On the Nature of the Affinities Connecting Organic Beings
      3. Morphology
      4. Development and Embryology
      5. Rudimentary, Atrophied, and Aborted Organs
      6. Summary
    15. XV: Recapitulation and Conclusion
  7. Glossary of the Principal Scientific Terms Used in the Present Volume
  8. List of Illustrations
  9. Endnotes
  10. Colophon
  11. Uncopyright

Endnotes

  1. Aristotle, in his Physicae Auscultationes (lib.2, cap.8, s.2), after remarking that rain does not fall in order to make the corn grow, any more than it falls to spoil the farmer’s corn when threshed out of doors, applies the same argument to organisation; and adds (as translated by Mr. Clair Grece, who first pointed out the passage to me), “So what hinders the different parts (of the body) from having this merely accidental relation in nature? as the teeth, for example, grow by necessity, the front ones sharp, adapted for dividing, and the grinders flat, and serviceable for masticating the food; since they were not made for the sake of this, but it was the result of accident. And in like manner as to other parts in which there appears to exist an adaptation to an end. Wheresoever, therefore, all things together (that is all the parts of one whole) happened like as if they were made for the sake of something, these were preserved, having been appropriately constituted by an internal spontaneity; and whatsoever things were not thus constituted, perished and still perish.” We here see the principle of natural selection shadowed forth, but how little Aristotle fully comprehended the principle, is shown by his remarks on the formation of the teeth. ↩︎

  2. I have taken the date of the first publication of Lamarck from Isidore Geoffroy Saint-Hilaire’s (Hist. Nat. Générale, tom. ii page 405, 1859) excellent history of opinion on this subject. In this work a full account is given of Buffon’s conclusions on the same subject. It is curious how largely my grandfather, Dr. Erasmus Darwin, anticipated the views and erroneous grounds of opinion of Lamarck in his “Zoonomia” (vol. i pages 500–510), published in 1794. According to Isid. Geoffroy there is no doubt that Goethe was an extreme partisan of similar views, as shown in the introduction to a work written in 1794 and 1795, but not published till long afterward; he has pointedly remarked (“Goethe als Naturforscher,” von Dr. Karl Meding, s. 34) that the future question for naturalists will be how, for instance, cattle got their horns and not for what they are used. It is rather a singular instance of the manner in which similar views arise at about the same time, that Goethe in Germany, Dr. Darwin in England, and Geoffroy Saint-Hilaire (as we shall immediately see) in France, came to the same conclusion on the origin of species, in the years 1794–5. ↩︎

  3. From references in Bronn’s Untersuchungen uber die Entwickelungs-Gesetze, it appears that the celebrated botanist and palaeontologist Unger published, in 1852, his belief that species undergo development and modification. Dalton, likewise, in Pander and Dalton’s work on Fossil Sloths, expressed, in 1821, a similar belief. Similar views have, as is well known, been maintained by Oken in his mystical Natur-Philosophie. From other references in Godron’s work Sur l’Espece, it seems that Bory St. Vincent, Burdach, Poiret and Fries, have all admitted that new species are continually being produced. I may add, that of the thirty-four authors named in this Historical Sketch, who believe in the modification of species, or at least disbelieve in separate acts of creation, twenty-seven have written on special branches of natural history or geology. ↩︎

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